Shark
The Greenland shark is the longest-living vertebrate. In 2016, radiocarbon dating aged one female at ~400 years; the species may reach 272–512. The secret is a metabolism crawling in the cold, deep Arctic. It takes ~150 years to mature — so one maturing today was born in the 1600s.
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Sharks have no real tongue — just a small slab of cartilage, the basihyal, on the mouth floor. Useless in most species, it is the cookiecutter shark's key weapon: the shark latches on, retracts its mobile basihyal to create a vacuum, then twists out a round plug of flesh.
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